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Nathan MillsNathan E. Mills, Ph.D. 
Associate Professor of Biology
Office: Pryor-England 161B
Phone: 501-279-4536


The Ecological Society of America

Partners in Amphibian and Reptile Conservation

All About Birds

Selected Publications

Teaching and Research Interests 

My general research interest is in understanding how the environment affects amphibian development and growth. My research has focused on two specific questions. First, how does hypoxia affect embryonic growth and development, and in turn, how does the embryo adaptively respond to hypoxia through changes in egg capsule oxygen conductance. I currently have undergraduate students involved in this research. And second, how do pesticides interact with environmental variables, such as competition and predation, to alter the survival and growth of amphibian larvae and the timing of metamorphosis. In addition to my primary research interest, I am involved in an amphibian survey at the Bald Knob National Wildlife Refuge near Bald Knob, Arkansas, and I have had the privilege of collaborating with Mike Plummer on research investigating the behavioral ecology of turtles and snakes.

My teaching responsibilities at Harding University include Human Anatomy and Physiology, Ornithology, and Ecology. I am also involved in team teaching Developmental Biology.


Department of Biology

Campus Box 12251
Pryor-England Science Center 
Harding University
Searcy, AR 72149-12251

Phone: 501-279-4459
Fax: 501-279-4706